Author ORCID Identifier

https://orcid.org/0000-0002-9561-3955

Semester

Spring

Date of Graduation

2023

Document Type

Dissertation

Degree Type

PhD

College

Eberly College of Arts and Sciences

Department

Biology

Committee Chair

Stephen P. DiFazio

Committee Member

Vagner A. Benedito

Committee Member

Gancho T. Slavov

Committee Member

Jennifer Hawkins

Committee Member

Craig F. Barrett

Abstract

Dissecting the genetic architecture of a highly complex trait can be a challenging task and insight into the mechanistic role of these genetic elements do not always follow. Association genetics and quantitative trait loci (QTL) mapping techniques are used to identify key genetic elements associated with complex traits. However, in order to characterize the mechanistic role of the interaction among these genetic elements that results in ultimate trait features, it may be helpful to peer into intermediate molecular phenotypes such as gene expression.

In this dissertation work we have genetically mapped complex traits related to biomass and wood chemistry in P. trichocarpa (black cottonwood), as they represent key adaptive traits for this species and are key commercially traits related to biomass recalcitrance in view of its potential as a bioenergy feedstock. Since, formation of wood involves the interaction of many biological pathways related to managing internal resources as well as reacting to biotic and abiotic stress, we have characterized the transcriptome of the xylem to glean its major features and interacting molecular pathways involved. Tree improvement towards modified wood chemistry is under considerable inertia due to lack of clear evidence on trade-offs related to biomass and stress tolerance. This work characterized the extent of genetic control and genetic correlations among these traits that may provide breeders a perspective of what they can expect when using standing genetic variation to improve biomass recalcitrance.

We leveraged a large multiparent controlled cross of P. trichocarpa to construct multiple genetic linkage maps, since accurate and dense maps are the cornerstone of successfully mapping traits that we examined in this study. We produced fine-scale recombination rate estimates and characterized its variation within the species and the P. trichocarpagenome. The maps produced and fine-scale recombination rates estimated here will be highly valued for future efforts to accelerate domestication of this species, as well as future studies that investigate broader questions such as evolutionary history, perennial development related to phenology, vegetative propagation, and dioecy that cannot be studied using conventional plant model systems such as Arabidopsis, rice, or maize. The genes or genomic regions we identified in this study together with the transcriptomic data we analyzed will compose an integral part of genomic selection and multi-omic predictive models that we aim to develop for this species

Comments

The dissertation has a companion Microsoft Excel file named 'chapters_2_3_4_Supplementary_tables.xlsx' that contains all the supplementary tables

Chapters_2_3_4_Supplementary_tables.xlsx (10997 kB)
Supplementary data tables

Share

COinS